A technique for efficient sorting of live termites from debris.
Psyche 100:1-4, 1993.
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PSYCHE
Vol. 100 1993 Nos. 1-2
A TECHNIQUE FOR EFFICIENT SORTING OF
LIVE TERMITES FROM DEBRIS
BY BARBARA L. THORNE AND NANCY L. BREISCH Department of Entomology
University of Maryland
College Park, MD 20742-5575
Sorting live termites from nest debris or soil is normally a labor- intensive procedure. Heavily populated nest sections or pieces of infested wood can be tapped over a collection dish, yielding satis- fying results at first but often diminishing rewards as termites hold on tenaciously or retreat to central cavities. Further, the process of tapping invariably also produces small pieces of debris as contami- nants in the collection tray. Those particles are unacceptable for clean samples for such purposes as reference collections, chemical or DNA analyses, weights, or volumetric samples. In these circum- stances, a second stage 'clean sort' is necessary. Among the techniques for picking out termites one by one, featherweight forceps are effective if administered gently. Some people prefer to use aspirators, but these should be limited to reverse-flow aspirators because of the hazard of lung infection from inhaling soil microbes or fungal spores present amidst termite debris. The receiving vial of an aspirator must be emptied fre- quently to avoid insect damage due to crowding. Soldier termites with defense secretions may obstruct aspirator suction tubes. Termitologists with sufficient patience use small paintbrushes to separate termites, nudging them towards a target receptacle or Manuscript received 12 February 1993.
Pachc 100: 1-4 11993). http //psyck mlclub.me/IOQ'100-001 hlml
2 Psyche [VOI. 100
waiting for them to crawl onto the brush, then collecting them with a quick flick into a container. Narrow cardboard strips (or folded index cards) are used in the same manner. Termite biologists use a variety of techniques to separate larger numbers of termites from broken nest fragments, splintered wood or soil. 'Papering off' or 'carding' termites works well for many species (Gay et al. 1955, McMahan and Watson 1977). In this approach one places a piece of paper or an index card on an active section of debris, letting the termites crawl onto the paper. Many species respond well to vertical positioning of the paper surface. When the card accumulates a large number of exploring termites, it is moved over to a collecting dish and tapped or flicked to dislodge the insects. The procedure is repeated as desired for collecting or sampling. Corrugated cardboard or moist paper towels are standard substrate variants for this technique, depending on conditions and the species under study.
Sieves with moisture below (eg. a container of moist sand or fil- ter paper) and/or heat above (light bulb) can work well to encour- age termites to sort themselves from other substrates by crawling through the sieve screen and dropping to the container below. Cau- tion must be taken, however, to check the apparatus frequently to avoid dessication.
In studies in which entire nests have been fumigated, water flotation methods are used to separate dead termites from nest and particle debris (Wiegert 1970, Darlington 1984). Isolating large numbers of subterranean termites from wood and soil debris is particularly challenging. These insects are typically small and delicate, and mortality from handling can be substantial with standard methods of obtaining clean samples. We describe here a quick, effective and inexpensive technique for sorting live Reticulitermes from debris. We have tested the technique with R. flavipes (Kollar), R. virginicus (Banks) and R. hesperus Banks. This approach may well be generalized or adapted for use with other termite genera.
Remove termites from their host substrate by tapping occupied sections over a collection tray or flat container. Splinters of wood debris, termite galleries and soil will also rain into the collection
19931 Thorne & Breisch 3
arena. Large fragments of wood should be removed to reduce harborage for the termites. After this initial crude separation, lay lengths of flexible plastic tubing (3116' internal diameter; 114' out- side diameter) in the collection tray, positioned such that the open ends are accessible to termites and the tube lies relatively flat. Ter- mites frequently travel along the perimeter of containers, so pru- dent placement of tube ends intersects these routes (especially corners). Open ends are best cut at angles rather than straight to expose more surface area at the opening. Tube length is not impor- tant, but 10-20 cm sections are convenient. Termites will soon begin to crawl into the tubes, packing in tightly with immigration greatly exceeding emigration. Termites prefer tight spaces over open areas, and are thus cooperative in remaining in the tubes. Leave the termites to self-sort; depending on numbers and activity level an interval of 10-60 minutes should yield good results.
When a tube is filled or sufficiently occupied, carefully lift it over a second collecting vessel and gently tap out the termites. Alternatively, a longer length of collection tubing (eg. 1 meter) can be draped over the edge of a lab bench, with one or both ends posi- tioned in a new collection receptacle. Termites crawl out of the tubing into the container, eliminating the need for handling or dis- turbance. Replace the tubing for additional sampling. 'Condi- tioned' tubes (previously occupied by termites, or formerly used as transit connections between two containers of termites) will out- perform clean tubing, but ultimately even new tubes will entice ter- mites.
The tube technique can be used in conjunction with other meth- ods; for example lengths of tubing placed in the collection recepta- cle under a sieve or Berlese sorting apparatus will attract dislodged termites and thus reduce dessication stress. The tube technique has a number of advantages over many other methods to collect Reticulitermes. After a quick, crude initial sepa- ration the biologist need return only occasionally to retrieve groups of uninjured, protected, debris-free specimens. DARLINGTON, J. P. E. C.
1984. A method for sampling the populations of large termite nests. Ann. Appl. Biol. 104: 427-436.
4 Psyche [vo~. 100
GAY, F. J., GREAVES, T., HOLDAWAY, F. G., WEATHERLY, A. H. 1955. Standard laboratory colonies of termites for evaluating the resistance of timber, timber preservatives, and other materials to termite attack. CSIRO, Bulletin #227, Melbourne, 60 pp.
MCMAHAN, E. A., WATSON, J. A. L.
1977. The effect of separation by 'papering' on caste ratios in Nasutitermes exitiosus (Hill) (Isoptera). J. Aust. Ent. Soc. 16: 455-457. WIEGERT, R. G.
1970. Energetics of the nest-building termite, Nasutitermes costalis (Holm- gren), in a Puerto Rican forest. In: Odum, H. T. A Tropical Rain Forest. Division of Technical Information, U. S. Atomic Energy Commission. pp. 57-64.
Vol. 100 1993 Nos. 1-2
A TECHNIQUE FOR EFFICIENT SORTING OF
LIVE TERMITES FROM DEBRIS
BY BARBARA L. THORNE AND NANCY L. BREISCH Department of Entomology
University of Maryland
College Park, MD 20742-5575
Sorting live termites from nest debris or soil is normally a labor- intensive procedure. Heavily populated nest sections or pieces of infested wood can be tapped over a collection dish, yielding satis- fying results at first but often diminishing rewards as termites hold on tenaciously or retreat to central cavities. Further, the process of tapping invariably also produces small pieces of debris as contami- nants in the collection tray. Those particles are unacceptable for clean samples for such purposes as reference collections, chemical or DNA analyses, weights, or volumetric samples. In these circum- stances, a second stage 'clean sort' is necessary. Among the techniques for picking out termites one by one, featherweight forceps are effective if administered gently. Some people prefer to use aspirators, but these should be limited to reverse-flow aspirators because of the hazard of lung infection from inhaling soil microbes or fungal spores present amidst termite debris. The receiving vial of an aspirator must be emptied fre- quently to avoid insect damage due to crowding. Soldier termites with defense secretions may obstruct aspirator suction tubes. Termitologists with sufficient patience use small paintbrushes to separate termites, nudging them towards a target receptacle or Manuscript received 12 February 1993.
Pachc 100: 1-4 11993). http //psyck mlclub.me/IOQ'100-001 hlml
2 Psyche [VOI. 100
waiting for them to crawl onto the brush, then collecting them with a quick flick into a container. Narrow cardboard strips (or folded index cards) are used in the same manner. Termite biologists use a variety of techniques to separate larger numbers of termites from broken nest fragments, splintered wood or soil. 'Papering off' or 'carding' termites works well for many species (Gay et al. 1955, McMahan and Watson 1977). In this approach one places a piece of paper or an index card on an active section of debris, letting the termites crawl onto the paper. Many species respond well to vertical positioning of the paper surface. When the card accumulates a large number of exploring termites, it is moved over to a collecting dish and tapped or flicked to dislodge the insects. The procedure is repeated as desired for collecting or sampling. Corrugated cardboard or moist paper towels are standard substrate variants for this technique, depending on conditions and the species under study.
Sieves with moisture below (eg. a container of moist sand or fil- ter paper) and/or heat above (light bulb) can work well to encour- age termites to sort themselves from other substrates by crawling through the sieve screen and dropping to the container below. Cau- tion must be taken, however, to check the apparatus frequently to avoid dessication.
In studies in which entire nests have been fumigated, water flotation methods are used to separate dead termites from nest and particle debris (Wiegert 1970, Darlington 1984). Isolating large numbers of subterranean termites from wood and soil debris is particularly challenging. These insects are typically small and delicate, and mortality from handling can be substantial with standard methods of obtaining clean samples. We describe here a quick, effective and inexpensive technique for sorting live Reticulitermes from debris. We have tested the technique with R. flavipes (Kollar), R. virginicus (Banks) and R. hesperus Banks. This approach may well be generalized or adapted for use with other termite genera.
Remove termites from their host substrate by tapping occupied sections over a collection tray or flat container. Splinters of wood debris, termite galleries and soil will also rain into the collection
19931 Thorne & Breisch 3
arena. Large fragments of wood should be removed to reduce harborage for the termites. After this initial crude separation, lay lengths of flexible plastic tubing (3116' internal diameter; 114' out- side diameter) in the collection tray, positioned such that the open ends are accessible to termites and the tube lies relatively flat. Ter- mites frequently travel along the perimeter of containers, so pru- dent placement of tube ends intersects these routes (especially corners). Open ends are best cut at angles rather than straight to expose more surface area at the opening. Tube length is not impor- tant, but 10-20 cm sections are convenient. Termites will soon begin to crawl into the tubes, packing in tightly with immigration greatly exceeding emigration. Termites prefer tight spaces over open areas, and are thus cooperative in remaining in the tubes. Leave the termites to self-sort; depending on numbers and activity level an interval of 10-60 minutes should yield good results.
When a tube is filled or sufficiently occupied, carefully lift it over a second collecting vessel and gently tap out the termites. Alternatively, a longer length of collection tubing (eg. 1 meter) can be draped over the edge of a lab bench, with one or both ends posi- tioned in a new collection receptacle. Termites crawl out of the tubing into the container, eliminating the need for handling or dis- turbance. Replace the tubing for additional sampling. 'Condi- tioned' tubes (previously occupied by termites, or formerly used as transit connections between two containers of termites) will out- perform clean tubing, but ultimately even new tubes will entice ter- mites.
The tube technique can be used in conjunction with other meth- ods; for example lengths of tubing placed in the collection recepta- cle under a sieve or Berlese sorting apparatus will attract dislodged termites and thus reduce dessication stress. The tube technique has a number of advantages over many other methods to collect Reticulitermes. After a quick, crude initial sepa- ration the biologist need return only occasionally to retrieve groups of uninjured, protected, debris-free specimens. DARLINGTON, J. P. E. C.
1984. A method for sampling the populations of large termite nests. Ann. Appl. Biol. 104: 427-436.
4 Psyche [vo~. 100
GAY, F. J., GREAVES, T., HOLDAWAY, F. G., WEATHERLY, A. H. 1955. Standard laboratory colonies of termites for evaluating the resistance of timber, timber preservatives, and other materials to termite attack. CSIRO, Bulletin #227, Melbourne, 60 pp.
MCMAHAN, E. A., WATSON, J. A. L.
1977. The effect of separation by 'papering' on caste ratios in Nasutitermes exitiosus (Hill) (Isoptera). J. Aust. Ent. Soc. 16: 455-457. WIEGERT, R. G.
1970. Energetics of the nest-building termite, Nasutitermes costalis (Holm- gren), in a Puerto Rican forest. In: Odum, H. T. A Tropical Rain Forest. Division of Technical Information, U. S. Atomic Energy Commission. pp. 57-64.
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Function Loops The Legend Of Psytrance WAV MiDi
Gold Class Audio Absolute Tech House WAV
Gold Class Audio Amazing Tropical House Vol 3 WAV MiDi
Gold Class Audio Future Boss WAV MiDi FL STUDiO XFER RECORDS SERUM
Gold Class Audio King Of House WAV MiDi
Gold Class Audio Main Progressive House WAV MiDi LD SYLENTH1 XFER SERUM
Gold Class Audio Major Pop 2 WAV MiDi
Haezer Analogue Onslaught WAV
Laniakea Sounds Atmospherica 2 WAV MiDi
Laniakea Sounds Aura WAV REVEAL SOUND SPiRE
Laniakea Sounds Best Of 2017 Ultimate Trance Collection WAV MiDi REVEAL SOUND SPiRE
Laniakea Sounds Downtempo Vibes WAV MiDi
Laniakea Sounds Future Bass Wave WAV MiDi LENNAR DiGiTAL SYLENTH1 REVEAL SOUND SPiRE
Laniakea Sounds Future Deep WAV MiDi
Laniakea Sounds Melodic Deep House Vol 2 WAV MiDi NATiVE iNSTRUMENTS MASSiVE
Laniakea Sounds Melodic Progressive WAV MiDi REVEAL SOUND SPiRE
Laniakea Sounds Mesmerizing Tech House Vol 2 WAV MiDi
Laniakea Sounds Mesmerizing Tech House Vol 3 WAV MiDi REVEAL SOUND SPiRE
Laniakea Sounds Modern Pop WAV MiDi REVEAL SOUND SPiRE
Laniakea Sounds Modern Vocal Loops 2 WAV
Laniakea Sounds Progressia WAV MiDi
Laniakea Sounds Progressive Chill WAV MiDi
Laniakea Sounds Progressive Vocal Chops 3 WAV
Laniakea Sounds Progressive Vocal Chops 4 WAV
Laniakea Sounds Tropicality WAV
Laniakea Sounds Vocal Atmospheres 3 WAV
Mayeniac Cloud God 2 Drum Kit WAV
Mayeniac DMSN Drum Kit WAV
Myloops Psy Trance Collection Sample Pack WAV and Studio One Template
Origin Sound Descend WAV MiDi
Origin Sound Elevate WAV MiDi
Origin Sound Monolith WAV MiDi
Origin Sound Revelation WAV MiDi
Production Master Berlin Techno WAV
Production Master Deep Tech House WAV MiDi
Production Master Downtempo Chill WAV
Production Master Future House Anthems WAV MiDi NATiVE iNSTRUMENTS MASSiVE LENNAR DiGiTAL SYLENTH1
Production Master Lavish Future Bass WAV
Production Master Lit Drill And Trap WAV MiDi
Production Master Miami 1982 WAV
Production Master Neurosys 2 WAV
Production Master Riddim Dubstep Weapons WAV ABLETON LiVE TEMPLATE XFER RECORDS SERUM
Production Master Stu Cause4Concern Trauma WAV XFER RECORDS SERUM
Production Master Summer Vibes WAV MiDi
Retrohandz Essential Moombah WAV
Splice Sounds Chef Pasquale Chef Szn Drumkit WAV
Splice Sounds Hiko Momoji Bedroom Sounds WAV
Splice Sounds Junkie Kid We Like Hard Sounds Vol 2 WAV MIDI FXP
TheBeatPlug Good Morning Daddy Drum Kit WAV
Tunecraft Sounds Ambient Cinematic Pads Vol 2 WAV MiDi XFER RECORDS SERUM
Tunecraft Sounds Future Bass Essentials Vol 2 WAV MiDi XFER RECORDS SERUM ABLETON LiVE PROJECT
Tunecraft Sounds Future Chill Hop Sessions WAV MiDi XFER RECORDS SERUM NATiVE iNSTRUMENTS MASSiVE
Tunecraft Sounds Twisted Tech Tools WAV NATiVE iNSTRUMENTS MASSiVE
WavSupply Josh Petruccio Godfather Drum Kit WAV